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An ion storm / herd of planet eating goats / Chup plague which moves round
the map, causing so much disruption it impels locals to migrate
or die.
Example: one theory about why the Roman Empire fell is that
the Mongol Hordes pushed the Huns out of their old lands. The Huns in turn
headed into the Empire.
As a scenario driver, a wave of something Bad
moving into the galaxy will force huge migrations and wars as people scramble
to escape its path.